Dr. Alfredo Hernández Llamas
[ Foto Personal ]

Doctor en Ciencias

CATEGORÍA: Investigador Titular C (ITC)



  • Programa de Acuicultura


  • Bioeconomía y sistemas expertos (SE) para el manejo acuícola
  • Nutrición Acuícola


  • Javier M. J. Ruiz-Velazco, Margarita Estrada-Perez, Nallely Estrada-Perez, Alfredo Hernandez-Llamas. 2022. Estimating the impact of COVID-19 pandemic on alternative semi-intensive shrimp (Penaeus vannamei) production schedules in Mexico: a stochastic bioeconomic approach. Aquaculture International https://doi.org/10.1007/s10499-022-00951-2
  • Javier M.J. Ruiz-Velazco, Miguel A. González-Romero, Nallely Estrada-Perez, Alfredo Hernandez-Llamas. 2021. Evaluating partial harvesting strategies for whiteleg shrimp Litopenaeus (Penaeus) vannamei semi-intensive commercial production: Profitability, uncertainty, and economic risk. Aquaculture International https://doi.org/10.1007/s10499-021-00695-5.
  • Nallely Estrada-Perez, Javier M.J. Ruiz-Velazco, Alfredo Hernandez-Llamas. 2020. Economic risk scenarios for semi-intensive production of Litopenaeus (Penaeus) vannamei shrimp affected by acute hepatopancreatic necrosis disease. Aquaculture Reports 18, 100442. https://doi.org/10.1016/j.aqrep.2020.100442
  • Nallely Estrada-Perez, Javier M. J. Ruiz-Velazco, Francisco J. Magallon-Barajas, Angel I. Campa-Cordova, Alfredo Hernández-Llamas. 2020. Dynamic stock model for analysing semi-intensive production of whiteleg shrimp Litopenaeus (Penaeus) vannamei affected by the acute hepatopancreatic necrosis disease: Assessment of disease severity indicators and relationships with pond water quality parameters. Aquaculture Research; 51:242-254. DOI: 10.1111/are.14370
  • Hernández-Llamas, A., L. Núñez-Amao, J. Naranjo-Páramo, M. Vargas-Mendieta and H. Villarreal-Colmenares. 2019. A review of alternative pond harvesting schedules for male and female redclaw crayfish Cherax quadricarinatus, reared in intensive monosex commercial conditions: a stochastic bioeconomic approach. Reviews in Aquaculture. doi: 10.1111/raq.12343
  • Estrada-Pérez, N., A. Hernández-Llamas, J.M.J. Ruiz-Velazco, I. Zavala-Leal, C.A. Romero-Bañuelos, E. Cruz-Crespo, C. Juárez-Rossete, D. Domínguez-Ojeda, A. Campos-Mendoza. 2018. Stochastic modelling of aquaponic production of tilapia (Oreochromis niloticus) with lettuce (Lactuca sativa) and cucumber (Cucumis sativus). Aquac Res. 2018;00:1–12. https://doi.org/10.1111/are.13840
  • Moreno-Figueroa L.D, Villarreal-Colmenares H., Naranjo-Paramo J., Vargas-Mendieta M., Mercier L., Casillas-Hernandez R., Hernandez-Llamas A. 2018. Bioeconomic modelling of the intensive production of whiteleg shrimp (Litopenaeus vannamei) in a photo-heterotrophic hypersaline system, with minimal seawater replacement. Reviews in Aquaculture, doi: 10.1111/raq.12252.
  • Moreno-Figueroa, L.D., J. Naranjo-Páramo, A. Hernández-Llamas, M. Vargas-Mendieta, J.A. Hernández-Gurrola, H. Villarreal-Colmenares. 2018. Performance of a photo-heterotrophic, hypersaline system for intensive cultivation of white leg shrimp (Litopenaeus vannamei) with minimal water replacement in lined ponds using a stochastic approach. Aquaculture Research 49:57-67. DOI: 10.1111/are.13432.
  • González-Romero, M.A., J.M.J. Ruiz-Velasco, M. Estrada-Pérez, J.T. Nieto-Navarro, I. Zavala-Leal, A. Hernández-Llamas. 2018. Assessing uncertainty of semi-intensive production of whiteleg shrimp (Litopenaeus vannamei) using partial harvesting programs. Aquaculture Research 49(2): 953-962.DOI: 10.1111/are.13542.
  • Naranjo-Páramo, J., A. Hernández-Llamas, M. Vargas-Mendieta, L. Mercier, H. Villarreal. 2017. Dynamics of commercial size interval populations of female redclaw crayfish (Cherax quadricarinatus) reared in gravel-lined ponds: A stochastic approach. Aquaculture. 484: 82-89.
  • Hernandez-Llamas A., Cabanillas-Ramos J., F.J. Magallon-Barajas. 2016. Estimating impact of white spot disease on economic risk in semi-intensive shrimp farms in Mexico: the case of the State of Sinaloa. Reviews in Aquaculture 8, 111–120. doi: 10.1111/raq.12084. ISSN: 1753-5123.
  • Estrada-Pérez, A., J.M.J. Ruiz-Velasco, A. Hernández-Llamas, I. Zavala-Leal, L. Martínez-Cárdenas. 2016. Deterministic and stochastic models for analysis of partial harvesting strategies and improvement of intensive commercial production of whiteleg shrimp (Litopenaeus vannamei). Aquacult Eng., 70: 56-62.Doi: 10.1016/j.aquaeng.2015.11.003.ISSN: 0144-8609
  • Hernández-Llamas, A. 2016. Stochastic assessment of economic losses associated with hurricane hazard for whiteleg shrimp Litopenaeus vannameicultivated in floating cages in northwestern Mexico. Aquac Res 47, 3359-3362.DOI: 10.1111/are. ISSN: 1355-557X
  • Núñez-Amao, L., H. Villarreal, J. Naranjo-Páramo and A. Hernández-Llamas. 2016. Stochastic analysis of production dynamics of male and female redclaw crayfish (Cherax quadricarinatus) reared under commercial intensive cultivation. Rev Aquacult. En prensa doi: 10.1111/raq.12170.
  • Hernandez-Llamas A., Magallon-Barajas F.J., Perez-Enriquez R., Cabanillas-Ramos J., Esparza-Leal H.M., Portillo-Clark G. 2014. Pond shutdown as a strategy for preventing outbreaks of the white spot disease in shrimp farms in Mexico. Reviews in Aquaculture, 6: 67- 74.
  • González-Romero, M.A., A. Hernández-Llamas, J.M.J. Ruiz-Velazco, T.N. Plascencia-Cuevas, J.T. Nieto-Navarro. 2014. Stochastic bio-economic optimization of pond size for intensive commercial production of whiteleg shrimp Litopenaeus vannamei. Aquaculture. 433:496-503.
  • Ruiz-Velazco, J.M.J., A. Hernández-Llamas and V.M. Gomez-Muñoz. 2014. A continuous diphasic model for prediction of survival of cultivated populations when affected by disease: the case of shrimp and white spot disease. Aquac Res. In Press DOI 10.1111/are.12454.
  • Hernández-Llamas, A., J.M.J. Ruiz-Velazco and V.M. Gomez-Muñoz. 2013. Economic risk associated with white spot disease and stochastic variability in economic, zootechnical and water quality parameters for intensive production of Litopenaeus vannamei. Reviews in Aquaculture 5: 121-131.
  • Ruiz-Velazco J.M.J., Estrada-Pérez M., Hernández-Llamas A., Nieto-Navarro J.T., Peña-Messina, E. 2013. Stock model and multivariate analysis for prediction of semi-intensive production of shrimp Litopenaeus vannamei as a function of water quality and management variables: A stochastic approach. Aquacultural Engineering 56: 34-41.
  • Hernandez-Llamas, A., Ruiz-Velazco J.M., Gomez-Muñoz, V.M. 2011. A stochastic approach for analysis of the influence of white spot disease, zootechnical parameters, water quality, and management factors on the variability of production of shrimp Litopenaeus vannamei cultivated under intensive commercial conditions. Aquacultural Engineering 45: 66-73. DOI: 10.1016/j.aquaeng.2011.07.002.
  • Hernandez-Llamas, A. and M. Zarain-Herzberg. 2011. Bioeconomic modeling and risk analysis of raising shrimp Litopenaeus vannamei in floating cages in northwestern Mexico: Assessment of hurricane hazard, stochastic variability of shrimp and feed prices, and zootechnical parameters. Aquaculture 314:261-268.
  • Ruiz-Velazco J.M.J, Hernández-Llamas A., Gomez-Muñoz V.M., Magallon F.J. 2010. Dynamics of intensive production of shrimp Litopenaeus vannamei affected by white spot disease, Aquaculture 300(1-4): 113-119.
  • Ruiz-Velazco J.M.J., A. Hernandez-Llamas, V.M. Gomez-Muñoz. 2010. Management of stocking density, pond size, starting time of aeration, and duration of cultivation for intensive commercial production of shrimp Litopenaeus vannamei. Aquacultural Engineering 43(3):114-119.
  • Zarain-Herzberg M., Fraga I., Hernandez-Llamas A. 2010. Advances in intensifying the cultivation of the shrimp Litopenaeus vannamei in floating cages. Aquaculture 300(1-4): 87-92.
  • Hernández-Llamas, A. 2009. Conventional and alternative dose-response models to estimate nutrient requirements of aquaculture species. Aquaculture. 292(4):207-213
  • Serviere-Zaragoza, E., A. Mazariegos-Villarreal, T. Reynoso-Granados, R. Robles-Hernández, P. Monsalvo-Spencer, G. Ponce-Díaz, A. Castro and A. Hernández-Llamas. 2009. Patterns of growth of juvenile pink abalone Haliotis corrugate fed re-hydrated natural feeds at a laboratory and a hatchery. Aquaculture Nutrition. 15(3):241-246.
  • Osuna-García, M., Hernandez-Llamas, A. and Mazón-Suástegui, J.M. 2008. Production dynamics of the giant lions-paw scallop Nodipecten subnodosus cultivated off-bottom. Aquaculture 264:260-267.
  • Alvarez, J.S., A. Hernández-Llamas, J. Galindo, I. Fraga, T. García, and H. Villarreal. 2007. Substitution of fishmeal with soybean meal in practical diets for juvenile white shrimp Litopenaeus schmitti (Pérez-Farfante & Kensley 1997). Aquaculture Research. 38:689-695.
  • Hernández-Llamas A., Balart E.F., Ponce-Díaz G., Civera-Cerecedo R. 2006. Feasibility of a new fishery in Baja California, Mexico based on the red crab Pleuroncodes planipes (Stimpson): Preliminary economic evaluation and risk assessment. Aquatic Living Resources 19:173-179
  • Jaime-Ceballos, B.J., A. Hernández-Llamas, T. García-Galano, and H. Villarreal. 2006. Substitution of Chaetoceros muelleri by Spirulina platensis meal in diets for Litopenaeus schmitti larvae. Aquaculture. 260:215-220.
  • Humberto Villarreal, Alfredo Hernandez-Llamas. 2005. Influence of temperature on larval development of Pacific brown shrimp Farfantepenaeus californiensis. Aquaculture 249: 257-263.
  • Hernandez-Llamas A. and D.A. Ratkowsky. 2004. Growth of fishes, crustaceans and molluscs: estimation of the von Bertalanffy, Logistic, Gompertz and Richards curves and a new growth model. Marine Ecology Progress Series 282: 237-244.
  • Hernandez-Llamas A., Gonzalez-Becerril A., Hernandez-Vazquez S. and S. Escutia-Zuñiga. 2004. Bioeconomic analysis of intensive production of the blue shrimp Litopenaeus stylirostris (Stimpson). Aquaculture Research 35:103-111.



  • Nallely Estrada Pérez, Doctorado, 2021: Bioeconomía del cultivo semi-intensivo de Litopenaeus (Penaeus) vannamei afectado por la necrosis aguda del hepatopáncreas: identificación y evaluación de factores de riesgo. Universidad Autónoma de Nayarit.
  • Lavinia Núñez Amao, Doctorado, 2018: Modelado bioeconómico de un sistema de producción intensiva de langosta de agua dulce Cherax quadricarinatus. Programa Interinstitucional en Bioeconomía Pesquera y Acuícola. CIBNOR, CICIMAR-IPN, Universidad Marista.
  • Luis Daniel Moreno Figueroa, Doctorado, 2017: Cultivo intensivo foto-heterotrófico del camarón blanco Litopenaeus vannamei (Boone, 1931) en alta salinidad con reposición mínima de agua. Centro de Investigaciones Biológicas del Noroeste.
  • Miguel Ángel González Romero, Doctorado, 2017: Modelo bioeconómico para el análisis de riesgo de la producción semi-intensiva de Litopenaeus vannamei con esquema de precosechas. Universidad Autónoma de Nayarit.
  • Margarita Estrada Pérez, Doctorado, 2016: Modelo bioeconómico para el análisis de riesgo de la producción semi-intensiva de camarón Litopenaeus vannamei.
  • Javier Marcial Ruiz Velazco Arce, Doctorado, 2011: Modelo Bioeconómico para el Análisis de riesgo del Cultivo Intensivo de Camarón Blanco (Litopenaeus vannamei). Mención honorífica. Premio Mejor Tesis de Posgrado 2011, Instituto Politécnico Nacional. Medalla Nayarit a la Investigación Científica y Tecnológica 2011.
  • Alfredo González Becerril, Doctorado, 2002: Modelación Bioeconómica de un Sistema de Producción de Camarón Litopenaeus stylirostris en Escuinapa, Sin. Mexico.


  • Nallely Estrada Pérez, Maestría, 2016: Modelos de producción de tres sistemas de recirculación agroacuícolas para el cultivo de tilapia, camarón y lechuga. Universidad Autónoma de Nayarit.
  • Alicia Estrada Pérez, Maestría, 2015: Modelo estocástico para el análisis de la variabilidad de la producción comercial intensiva de Litopenaeus vannamei con esquema de precosechas. Mención Honorífica. Universidad Autónoma de Nayarit.
  • Miguel Ángel González Romero, Maestría, 2013: Modelo bioeconómico para el diseño óptimo de granjas para la producción comercial intensiva de camarón blanco Litopenaeus vannamei: dimensión de los estanques. (Mención Honorífica). Universidad Autónoma de Nayarit.
  • Margarita Estrada Pérez, Maestría, 2012: Modelo bioeconómico de la producción semiintensiva de camarón blanco (Litopenaeus vannamei). Mención honorífica. Universidad Autónoma de Nayarit.


  • José L. Hernández y Martín González, Licenciatura, 1989: Rendimientos de producción de camarón azul Penaeus stylirostris a diferentes densidades de cultivo semiintensivo en Puerto Chale B.C.S. Universidad Autónoma de Baja California Sur.